I don't like the Nether forests because they represent a departure from what the Nether is meant to be, but I can see how fungi can be appropriate in a way because, in the real world, it breaks down materials (in this case, flesh).
With that thought in mind, the Blightlands is a biome of rot and decay. As opposed to netherrack or blackstone, its floors are littered mainly with rotting flesh--a renewable block which can be crafted with 9 rotten flesh items by the player, and replaces netherrack as the block that crimson nylium grows on. Some features include boulders made of either netherrack or blackstone (which can contain the new gold ores), some small bone spires/pillars which jut out of the ground, walls, and ceilings with flesh still clinging to parts of them, and the crimson forests we have now.
Sometimes this biome will be completely covered by crimson forest, and sometimes parts of it will lay bare to fester in the open. Nylium still won't spread on its own, and will require the player to bonemeal adjacent flesh blocks. Alternatively, rotten flesh could replace bonemeal for growing crimson biota as a whole.
Axes are the appropriate tool to "mine" rotting flesh blocks.
Edit: What if leaving rotting flesh blocks in the soul sand valley eventually turned them into soul soil?
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